Bernhard Zipfel

Carol V. Ward

Brian G. Richmond

The 1924 African discovery of an early hominin child's skull, referred to as Australopithecus africanus by Raymond Dart, was a major event in the history of paleoanthropology. This provided the first evidence of early hominins in Africa and overturned conventional ideas about human evolution. Subsequent discoveries of A. africanus fossils, notably from cave deposits at Sterkfontein, yielded the first evidence that early hominins were habitual bipeds. Fifty years after this, the discovered wealth of fossil evidence in eastern Africa of the slightly older and craniodentally more primitive taxon, A. afarensis, catalyzed debates about the origin and evolution of human gait and the phylogentic relationships among early hominins. This formed the main basis of our understanding of early hominin bipedality and paleobiology. Little attention has been paid to the variation among species in postcranial anatomy and locomotion, although intriguing hints are beginning to appear in the literature. Did multiple varieties of bipedality evolve? Did australopith species differ in positional or manipulative abilities, body proportions, or patterns of sexual dimorphism? These are critical questions for understanding the evolution of australopiths and hominin locomotion. In this book, Bernhard Zipfel, Brian Richmond, Carol Ward, and the most knowledgeable scholars in their respective fields provide groundbreaking accounts for each postcranial fossil and expert examinations into the background of each fossil. The chapters include standardized high-quality photographs and anatomical descriptions to allow readers to read the book entirely or learn by comparing features across chapters. Hominin Postcranial Remains from Sterkfontein, South Africa, 1936-1995 is an evolutionary history of South African hominins, and it offers readers an orientation and introduction to the field. This is an important reference book for professional paleontologists, paleobiologists, anthropologists, geologists, students, and historians interested in human evolution.

Hominin Postcranial Remains from Sterkfontein, South Africa, 1936-1995

The Sterkfontein hominin fossils generally are attributed to the species Australopithecus africanus, because most craniodental remains from the site are attributable to that taxon (reviews in Grine, 2013, 2019). However, there may be more than one hominin represented within the sample, even within the most productive Member, Member 4, and given the complex stratigraphy of the site and challenges in dating the deposits, this may or not may be the case. In general. Several studies have suggested the presence of two or more australopith taxon within the sample, each citing more morphological variation among the craniodental remains from Sterkfontein that can be attributed to a single species, at least compared to extant hominoid taxa (Kimbel and White, 1988; Clarke 1988, 1994; Lockwood, 1997, Lockwood and Tobias, 2002). However, it is notable that none of these studies agree on which specimens comprise the different possible taxa or groups, largely due to emphasis on different aspects of morphology varying among the fossils. The likely time depth of the Sterkfontein sample, even within Member 4, (see Chapter 3, this volume), may also complicate assessment of potential taxonomic heterogeneity at the site. None of the Sterkfontein postcranial fossils can be definitively associated with any craniodental specimens (but see Thackeray et al., 2002), and so cannot be related directly to any of the proposed taxonomic divisions within the sample. However, some studies have cited variation within the postcranial fossils that may also reflect taxonomic variation, although many studies to date have not tackled this question rigorously. Even though these suggestions have been made occasionally in the literature, no clear or consistent suggestion of two or more taxa has been apparent within the postcranial samples (reviewed in Grine 2019). Taxonomic variation is one of the key questions that each chapter in this volume addresses (summarized in Chapter 18, this volume)"--

Bernhard Zipfel

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