Gustav Arthur Cooper

No studies in depth have been made of the brachiopods from the Jurassic deposits of Saudi Arabia. This first study of the brachiopods from this important region is based mainly on a collection presented to the Smithsonian Institution by the Arabian-American Oil Company (Aramco). In addition, the study includes collections made by Drs. P.M. Kier and E.G. Kauffman of the Smithsonian Institution and R.W. Powers, C.D. Redmond and H.A. MacClure of the Arabian-American Oil Company. Sixty-one genera are described of which 29 are new. Of these 13 rhynchonellid genera are new: Amydroptychus, Baeorhynchia, Colpotoria, Conarosia, Deltarynchia, Echyrosia, Eurysites, Heteromychus, Lirellarina, Nastosia, Pycnoria, Schizoria, and Strongyloria. Other described genera are: Burmirhynchia Buckman, 1917, Cymatorhynchia Buckman, 1917, Daghanirhynchia Muir-Wood, 1935, Gibbirhynchia Buckman, 1917, Globirhynchia Buckman, 1917, Kallirhynchia Buckman, 1917, Kutchirhynchia Buckman, 1917, Somalirhynchia Weir, 1925, Sphenorhynchia Buckman, 1917, and Torquirhynchia Childs, 1969. Of Spiriferinacea, one genus, Calyptoria, is new and two genera are described: Liospiriferina Rouselle, 1977, and Spiriferina d'Orbigny, 1847. The Terebratulacea are represented by 11 new genera: Arabatia, Arabicella, Arapsopleurum, Arapsothyris, Dissoria, Ectyphoria, Pionopleurum, Pleuraloma, Stenorina, Tanyothyris, and Toxonelasma. Seventeen described terebratulaceans are Apatecosia Cooper, 1983, Avonothyris? Buckman, 1917, Bihenithyris Muir-Wood, 1935, Dolichobrochus Cooper, 1983, Dorsoplicathyris? Almeras, 1971, Glyphisaria? Cooper, 1983, Gyrosina? Cooper, 1983, Habrobrochus Cooper, 1983, Kutchithyris? Buckman, 1917, Loboidothyris? Buckman, 1917, Orthotoma Quenstedt, 1869, Plectothyris? Buckman, 1917, Pseudowattonithyris? Almeras, 1971, Somalithyris Muir-Wood, 1935; Sphaeroidothyris Buckman, 1917, Stiphrothyris? Buckman, 1917, and Striithyris Muir-Wood, 1935. The Zeilleriacea include four new genera: Apothyris, Mycerosia, Sphriganaria and Xenorina. Described zeilleriids are Flabellothyris Eudes-Deslongchamps, 1884, Rugitela Muir-Wood, 1936, and Zeilleria Bayle, 1878. A total of 166 species are described and 25 lots are identifiable as species. Pseudoglossothyris? sulcata Muir-Wood, 1935, from Somaliland (Somali Republic) is shown to be a zeilleriid, and the species is herein transferred to Aulacothyris. Eudesia cardioides Douvill, 1916, is herein transferred to the new genus Sphriganaria. The Liassic Marrat Formation abounds in spiriferinids. The Dhruma Formation (Bajocian to Callovian) is rich in rhynchonellids which dwindle in numbers in late Dhruma beds. The overlying Tuwaiq Mountain and Hanifa formations (Callovian to Kimmeridgian) are conspicuous for the large numbers of terebratulaceans which far outnumber the rhynchonellids. Correlation with Jurassic sequences near and far is difficult because of the high degree of endemism shown by the Saudi Arabian brachiopods. Precise correlation with British and European faunas is not now possible. Relationships with the Jurassic faunas of the Sinai, Israel and East Africa in the Callovian is suggested by the presence of two species in common and generic representation shown by Daghanirhynchia, Somalirhynchia, Bihenithyris, Somalithyris, and Striithyris.

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